Lec 12 Full Delivery Script - Color Patterns I

2026-05-05 11:27:55 • 19:02

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Lecture 12, Color Patterns I.

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EB 187, Ecology and Evolution of Color Week 6 Professor Michael Alfaro May 7th, 2026

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Source Deck.

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LEC 12, Slides.QMD, 23 Content Slides Plus 1 Title, Q Conventions.

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For Fragment Reveals, For Think Pair Share, Stage Directions in.

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Title Slide.

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Good Morning.

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Lecture 12, Color Patterns.

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On Tuesday, we ended with the mixing continuum, the synthesis of the entire color mechanism

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unit.

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Today we leave mechanism behind and ask the next question.

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Once a mechanism is in place, how does the body decide where to put it?

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Slide 1, Tuesday recap.

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Six Photographs.

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Six positions on the continuum.

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Quick pass.

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Position 1, Turrico.

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Pure pigment.

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Turrico-Verdin in the Feather Barbe.

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Bleach the Feather, color is gone.

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Grind the Feather, color survives.

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Position 2, Tannager.

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Karatanoid in the Cortex Plus, an ordered keratin layer beneath.

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The reflective backing double saturation.

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Position 3, Papilio.

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Yellow pigment plus a concave multi-layer adding blue iridescence.

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I mixes them to emerald.

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Position 4, Green Parrot.

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Spungy beta keratin makes the blue.

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Cidico-Folven filters at yellow.

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Subtractive mixing in tissue gives green.

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Position 5, Peacock.

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Melanon rods packed in a 2D photonic lattice.

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The pigment is the building block of the structure.

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Position 6, Morpho.

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Christmas tree ridges of clear chiton.

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No pigment in the blue.

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Bleach does nothing.

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Grind destroys it.

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Mechanism is set.

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Today, how does it get arranged in space and stripes, bars and spots?

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Slide 2, Cuddlefish camouflage.

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Last week, we talked about chromatophores as units of color.

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Here is what they look like in action.

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Sephalopod chromatophores expand and contract in milliseconds.

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Same machinery as the Salopris and Tree Frog from Tuesday, just on a faster clock.

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Slide 3, Camelian iridophores.

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The video opens with a Morpho butterfly intro, a LEC-11 callback.

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The structural color framework.

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Then it cuts to the Camelian.

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Watch the iridophore lattice spacing change.

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Cells expand.

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Latis spacing increases.

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Reflected wavelength shifts red.

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That is structural color modulated by cell biology, pure LEC-9-10 physics, with LEC-11

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cellular control.

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Slide 4, Bird of Paradise.

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The superb bird of paradise.

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Watch the courtship display.

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This bird is using mechanism, structural color in the throat shield, super black background

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feathers from LEC-11 and a precisely arranged spatial pattern.

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The pattern is the signal.

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Three questions latent in this video.

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First, how does the male assemble those colors?

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LEC-8-11.

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We did that.

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Second, how does the body know to put those patches where they are?

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Today's lecture.

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Third, why does the female care?

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LEC-14-16.

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Coming up.

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Today's pivot is from mechanism to arrangement.

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Slide 5, three questions.

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The agenda for today.

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One, what patterns exist?

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Stripes, bars, spots, a cell-eye, reticulations.

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Two, where do the cells come from?

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Neural crust for chromata-4 lineages.

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Three, how do cells self-organize?

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Short-range repulsion.

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Long-range help.

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Turing dynamics.

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The arc note at the bottom.

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Mechanism.

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LEC-8-11.

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Arrangement today.

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In LEC-14-16, we are in the middle of a four-week structural arc.

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Slide 6, pattern diversity.

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Eight photographs.

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I'm not going to lecture.

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I'm going to point.

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Tiger.

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Zebra.

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Leopard.

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Giraffe.

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Cuddlefish.

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Dark frog.

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Bicyclus eye spots.

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Peacock eye spots.

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Every one of these started as an un-patterned embryo.

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What rules could possibly produce all of this?

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A single developmental program would be impossible.

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But a small toolkit of patterning rules applied to different cells with different parameters

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in different body regions can produce everything you see here.

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That is the through line for both days.

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Slide 7, naming patterns.

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A taxonomy of pattern elements.

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Stripe.

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Bar.

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Spot.

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Ocelas.

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Reticulation.

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Vermiculation.

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Don't memorize this.

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Use it as a reference.

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Two terminology warnings.

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Ictheologists call horizontal bands stripes.

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Some herpetologists call vertical bands stripes.

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The Nymphilid ground plan is the lepidoptuist system.

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Cross-taxon comparisons need explicit definitions.

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The important pedagogical move is here.

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Endler 1978 pattern is measurable.

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Spot size, spacing, contrast, orientation.

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Modern tools, microtoolbox, patTR3D do the same job

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in software.

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For the project, you should think about pattern in measurable terms, not just verbal categories.

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A pattern is only a pattern if a viewer can resolve it.

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LEC's six spatial acuity sets the pattern's effective grain.

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Slide 8, cell palette.

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The chromato-4 lineages.

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We met these in LEC-11 at the molecular level.

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Today we treat them as spatial agents, units that get placed somewhere in the skin.

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Melanophor, melanin, black and brown.

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LEC-8, Santa IV, Teradins and Caratenoids Yellow.

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LEC-8, Erithro IV, Keto Caratenoids Red.

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LEC-8, Iridiphor, Guanine platelets, Blue and Silver.

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LEC-9 and 10.

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Mammals and birds have only melanocytes.

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Pattern complexity reflects palette size.

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Fish and herps with the full four cell palette get an order of magnitude larger pattern phenotypes

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face.

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That asymmetry is going to come back when we hit Murray's Scaling role later.

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Slide 9, TPS, Cell Stack.

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Three organisms.

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Emperor Angelfish.

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Lepard.

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Dark frog.

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Pairs for each identify which chromato-4 types are involved and what stacking order.

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90 seconds.

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Talk to your neighbor.

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The expected answers.

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Emperor Angelfish.

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Full palette.

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Iridiphor, Guanine layer giving the blue.

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Melanophor's stripe pattern on top.

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Santa IV over yellow regions.

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Stacking matters for the green where Santa IV and Iridiphor overlap.

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Lepard.

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Melanocytes only.

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The spotted appearance comes entirely from spatial regulation of one cell type.

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You melanin clumps in spot regions.

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Dark frog.

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Three layer chromato-4 unit from LEC-11.

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The four on top.

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Iridiphor middle.

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Melanophor deep.

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Here is the reveal.

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The dark frog and the Emperor Angelfish use the same chromato-4 stack.

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The leopard uses one cell type.

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The cell palette doesn't predict the pattern.

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The rules of cell placement do.

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That is where Turing comes in.

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Slide 10, two ways.

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This is the conceptual heart of the lecture.

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Two genuinely different developmental logics.

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Both real.

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Both operating in nature.

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Three pattern.

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Wulpert 1969.

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The French flag model.

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A morphogen is set up across a tissue.

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Bicoid in flyambrias.

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BMP in vertebrae limbs.

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Cells read concentration like reading a thermometer and decide identity from where they sit.

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Identity is positional.

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Body coordinates map to color.

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Examples.

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Butterfly eye spots.

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Fly bristles.

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Vertebrae limb digits.

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Cell organization.

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During 1952, reaction diffusion.

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Two diffusible signals couple.

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Activator promotes itself plus the inhibitor.

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Inhibitor diffuses faster.

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Periodic instability emerges spontaneously.

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There is no pre-existing map.

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The pattern is the dynamics.

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Examples.

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Zebrafish stripes.

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Jaguar rosettes.

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Cichlid bars.

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Real animals do both.

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Different body regions can use different logic and the same animal.

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A leopard's body is a touring canvas.

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A leopard's face is positional information.

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Today we tour the evidence for each.

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Slide 11 condo angel fish.

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This is the single most persuasive slide in the developmental biology side of the course.

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Pomechanthus imparata.

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The emperor angel fish.

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As the fish grows, the number of stripes increases.

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If stripes were a fixed pre-pattern, they would just spread apart as the body lengthens.

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Instead, new stripes appear by branching.

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Exactly what a touring simulation predicts.

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Condo and Ause wrote down the equations and matched the stripes-facing dynamics quantitatively.

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This is the proof that touring operates in a real animal.

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Forty-three years after touring's 1952 paper proposed reaction diffusion.

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The biology lagged the math by half a century.

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Then the biology arrived.

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Slide 12 cheetah cubs.

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Cone is not a fixed image.

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It is a developmental trajectory.

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Cheetah cubs are born with a long-silvery dorsal mantle that obscures their spots.

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The mantle fades over the first three months.

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The adult phenotype emerges.

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Same point applies to taper babies, lion cubs, wild boar piglets.

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Stripes that fade with age.

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A simple touring parameter set is unlikely to explain this.

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There must be temporal modulation of the patterning machinery.

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The bridge to the previous slide is intentional.

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Condo just showed us pattern rearranging on a growing fish.

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Cheetah cubs show pattern fading on a growing mammal.

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Pattern is dynamic across the lifespan.

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Slide 13.

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Turing math.

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Minute Earth.

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The equations on screen are exactly Turing's 1952 reaction diffusion equations.

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The cartoon explainer just walked us through the math.

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Now let's see how those equations get extracted from real cells.

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Number 14.

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Cell rules.

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Nakamasu at all 2009 used cell ablation in zebrafish to extract the actual cell interaction

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rules.

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Remove melanophores and defined regions.

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Watch what happens to surrounding zanthophores.

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Two rules.

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Rule 1.

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Short range.

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Melanaphore and zanthophore repel each other.

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Rule 2.

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Long range.

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Melanaphore promotes zanthophore survival via gap junctions.

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Watanabe and Condo 2015 showed the long range channel is gap junction mediated.

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The fronthofer 2013 zebrafish mutant figure.

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Wild type at panel A full stripes.

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Panel C through F. Iridiform mutants RSE and Shud.

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Stripes break up into spots.

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Same Turing system.

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One gene perturbed.

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Spots not stripes.

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The leopard zebrafish story.

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Watanabe 2006 is the same logic with a different gene.

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The next in 41.8.

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The pedagogical point is not the math.

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It is this.

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Cells follow simple local rules.

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A single gene change alters the rules.

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The global pattern reorganizes.

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That is Turing flavor evo-devo in one slide.

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Slide 15.

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Cell migration video.

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Watch the cells move.

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Black cells are melanophores, moving by a me-boyd motility.

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Yellow cells are zanthophores, filling the spaces.

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They repel each other on contact.

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They sustain each other at distance through gap junctions.

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After hundreds of cell cell interactions, stripes emerge.

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There is no master plan.

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No coordinate system.

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Just thousands of cells executing two interaction rules.

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The pattern is what emerges when the rules run to equilibrium.

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LEC 11 callback.

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Last week we met chromatophores as the units of color.

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Today we are seeing them as the units of pattern.

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Cells 2 functional roles.

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Slide 16.

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Labyrinth intuition.

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Why a labyrinth?

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Every white wiggle is a high activator pocket.

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A region of cells locked into one state.

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Around each wiggle, a faster spreading inhibitor keeps neighbors at bay.

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Three local rules produce the labyrinth.

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One.

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Activator self promotes.

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Wiggle tips can extend.

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Two.

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Inhibitor diffuses faster.

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Wiggles can't merge.

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Three.

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Tips bend away from other tips.

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Wiggles snake and branch instead of straightening and stripes.

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The parameter knob.

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Tweak the diffusion ratio.

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Ratio up stripes.

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Middle labyrinth.

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Ratio down spots.

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Same equations.

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Three different patterns.

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No map.

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No designer.

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Just chemistry.

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This exact labyrinth zone is where every spotted by inverse spotted, salmoned hybrid

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lands.

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Once a parent at one end of parameter space with a parent at the other end.

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The F1 sits in the labyrinth zone in between.

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We will see it Tuesday.

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Slide 17.

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Murray scaling.

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A teaser.

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Tuesday we go deeper.

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Today the punchline.

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Mouse uniform.

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Body too small for one wavelength.

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Cheetah spots.

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Lepard rosettes.

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Tiger stripes.

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Elephant uniform.

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Too large.

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Body damps.

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Same touring equation.

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Body size is the boundary condition.

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James Murray showed this with a single equation in 1981.

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The bonus fact.

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Spotted cuts can have striped tails.

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But striped cuts never have spotted tails.

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Cheetahs have stripes on the tail tip.

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Tigers don't have spots.

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The model predicts this.

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At the small diameter of a tail, the same activator inhibitor dynamics produce stripes

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by default.

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LEC 11 callback.

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This is mostly a mammal story because mammals have only one chromatofore type.

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The same scaling rule applied to fish or herp would be cleaner if the cell palette weren't

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already adding pattern dimensions.

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Slide 18.

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I spots.

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The alternative logic to touring.

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Bicyclus and inana.

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Wing I spots.

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The focal cell.

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A small group of cells in the late wing disc.

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Secreta morphogen.

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Surrounding cells read the gradient and decide their identity from where they sit.

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Concentric pigment rings.

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That is positional information in the strict sense.

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The experimental punchline is striking.

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Breakfield, Carol.

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Bell-Dade transplant experiments.

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Take a fragment of larval wing disc containing the focal cells.

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Graphed it to a new location on another wing.

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An ectopic eye spot appears centered on the graft.

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The eye spot is positionally specified by the focal cells.

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The connection to mechanism layers is nontrivial.

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The rings of an eye spot are not monochromatic.

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The inner black ring is melanin LEC 8.

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The outer rings often involve structural color, LEC 9 and 10.

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So an eye spot is positional information saying where to put rings, plus mechanism saying

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how each ring colors itself.

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Both lecture stack.

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Same animal class as zebrafish.

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Completely different developmental logic.

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Slide 19, Mimulus.

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Plant petal patterns are nearly always walt-pert.

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Plants don't have migrating cells.

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Anthosine and biosynthesis is decided locally, positioned by position, based on transcription

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factor gradients of MYB, BCHLH and WD40 proteins.

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Animals migrate cells so they can self-organize.

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Plants don't migrate cells so they can't.

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The Mimulus Luicyye, the bumblebee pollinated monkey flower.

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The bullseye.

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In VUV vision, LEC 6 callback, the center is darker.

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UV absorbing anthosine.

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The periphery is brighter.

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The bullseye is invisible to humans, but obvious to a foraging bee.

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Pollinator vision sets the receiver context for plant pattern.

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Pattern plus receiver equals signal.

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That is next week.

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Slide 20 synthesis.

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!

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Turing.

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Local rules.

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Periodic output.

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Zebrafish, Sicklid, Jaguar, C-shell.

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Pre-pattern on the right.

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Walt-pert.

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Gradient.

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Positional readout.

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Butterfly icepot.

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Feather barbule.

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Vertibrate digit.

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Most real animals use both in different body regions.

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The same animal can be touring on the body and Walt-pert on the face.

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Slide 21.

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Project TPS.

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Pull up the fish photo you've chosen for the project.

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Three minutes with your neighbor.

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Three questions.

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One cell palette.

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Which chromatophore types are present?

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Eradessent blue means iridipore.

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Matt-black means melanophore dominant.

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Saturated red means erytherphore.

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Two, Patterning Rule.

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Does the pattern look more positional with specific markings that specific locations

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or self-organized with periodic stripes or spots?

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Three, one testable prediction.

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If you knocked out melanophore's anthropore communication, what would change?

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The project framing is now structured.

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Three questions.

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Every fish.

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Every time.

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Bring your answers to Lab Wednesday.

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They will seed the project right up.

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Slide 22.

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Looking ahead.

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Tuesday.

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Five threats.

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Pattern blending.

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Mia's always hybrid salmonates.

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Murray's Scaling Rule.

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Full version.

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With the Cheetah Tail prediction.

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Evodivo, Heliconius, Master Regulators, W and T.A. Optics, Double Sex.

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Single gene flips.

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Aguda, Leopard, ALX3.

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Bridge to function.

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Same patterns.

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Three different jobs.

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Entrance ticket Tuesday.

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Predict the F1 hybrid pattern from two salmated parents.

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Bring your intuition.

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We will test it.

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Slide 23.

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Exit ticket.

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Three options.

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Pick one.

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Could a butterfly icepot form by a turn dynamics?

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Why or why not?

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The could a zebrafish stripe form by a positional information?

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Why or why not?

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See Pick One organism from today and identify its cell palette and likely patterning rule.

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Submit on Bruin Learn before midnight.

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The framework you used in the project TPS is the framework you will use for the rest

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of the unit.

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Today we made the move from mechanism to arrangement.

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Tuesday we go from arrangement to evolution.

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See you Tuesday.